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Netherlands, the United Kingdom, Poland, Spain, France, Belgium, and Sweden (6,7,12,15,17,23,24). Susceptible nursery varieties include, but are not limited to: Catawbiense Grandiflorum, Catawbiense Boursalt, Schneewolk, Colonel Coen, Gomer Waterer, and Cunningham’s White. Extreme care should be exercised when handling and diluting the first round of PCR as DNA contamination is likely to occur at this stage.
We are grateful to the Gordon and Betty Moore Foundation, the USDA Forest Service Pacific Southwest Research Station, USDA Forest Service Forest Health Management, and the California Department of Forestry for providing funding to support the research underlying this Diagnostic Guide. Thin black zone lines characteristically separate diseased from healthy tissue in oak and tanoak cankers (B).

Several years after onset of infection, rapid browning and death of all leaves occurs in some oak trees infected with P.
Necrosis is often concentrated along the midvein in leaves infected from a twig infection via the petiole (A).
Leaves infected without prior twig infection can have leaf spots on any part of the leaf surface (B, C, and D).
The role of Phytophthora pathogens in forests and semi-natural communities in Europe and Africa. Garbelotto, Department of Environmental Science, Policy and Management, Ecosystem Science Division, 151 Hilgard Hall, University of California, Berkeley 94720; E.

Hansen, Department of Botany and Plant Pathology, Oregon State University, Corvallis 97331; D.

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