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To reach a series of caves dug into a cliff 155 feet above the valley floor, Matt Segal scales a rock face so fragile it often breaks off to the touch. On the shore of Canada's Lake Ontario, Zhang enjoyed watching the elegant feathers of mute swans. Mangroves contribute to the system by trapping reef-bound sediment, filtering out pollution, and serving as nursery for many reef fish and invertebrates. A view from 12,000 feet, off the coast of Belize, shows the parts of the system that make the whole.
Climbers and scientists follow a trail above the Kali Gandaki River in Nepal’s remote Mustang region.
Yachts bob in Botafogo Bay, cradled between the beach and the tall rock known as Sugar Loaf. A school of chubs swims over the coral reef of Cordelia Banks near the island of Roatan in Honduras.
As a medical student in Taichung, Cheng visited Gaomei wetland, a protected habitat where water sculpts the mud. In the late afternoon on Ipanema, beachgoers play the popular game of altinho, or keep it up, using soccer moves to pass the ball in the air from person to person.


Dusk falls over the temples and homes of Tsarang, once the region’s most important town.
Sea grass is not the typical meal for the primarily carnivorous species, which feeds on jellyfish, crabs, and conchs.
While feeding, this one opened its wings and gave the photographer the shot he'd hoped for. The arched roots of mangroves like these form gateways through which multitudes of juveniles swim toward adulthood on the reef.
Here, at the upper end of the Mesoamerican Reef, convene the largest known assemblies of whale sharks.
The reef’s calcium carbonate city teems with species, among them this spiny-headed blenny. Cordelia Banks, at the southeastern end of the Mesoamerican Reef, has the greatest abundance of corals known in the Caribbean. Scarus guacamaia, the largest herbivorous fish in the Atlantic, uses all three provinces of the Mesoamerican Reef in the course of its life. In Mustang, where the centuries have not disrupted the traditional rhythm of life, the caves offer clues to a time when the remote Himalayan kingdom was a hub linking Tibet to the rest of the world.


Next comes the white line of coral rubble along the reef crest, then the sandy back reef, and, finally, the lagoon: a maze of sand islets, mangrove cays, and sea grass beds. Researchers believe the banks, not fully protected, are a crucial nursery from which larval coral-reef organisms seed other regions of the Caribbean. As a juvenile it seeks the protection of the submerged roots at the fringe of the mangrove forest, from which it makes foraging runs to adjacent sea grass beds.
Scientists are helping sharks acquire a taste for the invaders by feeding them speared lionfish. Farther south, off the coast of Belize, whale sharks are drawn to the white clouds of eggs released by huge aggregations of spawning dog snappers, mutton snappers, and cubera snappers.



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