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Their service opened to the general public in 2009 after several years of conceptualization. 500px features work from thousands of photographers, and they make it easy to find some of the best work through voting algorithm and editors choice categories.
At the time of this writing there are 7 basic portfolio themes, and 4 additional premium themes reserved for paid subscribers.
I highly recommend that semi-professional to professional level photographers check out the site, and if they like what they see sign up for a free account. One of the major disadvantages of 500px is that the content isn’t indexed very well (as you pointed out). I maintain a pro level Flickr account, but rarely use the service for anything besides a way to publish a large amount of photos online for others to browse.
I don’t have much experience with SmugMug, but it appears to be a halfway point between Flickr and 500px. My best advice is that you start out with a free account and if you end up liking it after a couple weeks upgrade to a full account. Since that time they have gained nearly 50,000 members and currently receive around 1.4 million visits each month. Specifically they have created a social networking platform directed towards professional level photographers. When you follow another member you will see all of your friend’s most recent contributions displayed on your personalized homepage.
If another user comments on or favorites one of your photos, that action will show up in the activity feed. First you must agree to the store policy and verify that you own the rights to your photographs. If you are an avid photographer who has a large number of photographs that you want to share, then this feature would be essential. If you aren’t interested in building a wholly unique portfolio site then $50 is a pretty good deal for such a sleek interface.

Their service is slowly expanding with new features, and their web traffic is quickly growing.
Amateur photographers might be a little more discouraged from user feedback on the site, so I recommend that you limit your portfolio to your absolute best work. But both 500px and smugmug lacks good searching and grouping function at where Flickr still leads.
But I am afraid that I couldnt see the works of photographers with similar interest like me. Most work being uploaded isn’t getting tagged with enough keywords to have many images turning up in search results. It offers some of the same social networking features as 500px and also the more general purpose photo sharing features of Flickr. The free account has most of the same features and will give you a good idea of whether it’s worth your money to invest in a 1 year subscription. They have built a community that is filled with talented artists, which in return generates a large amount of traffic. Whenever you start following another photographer, or another photographer follows you, that will also show in your feed. I don’t fully understand the exact methods involved in determining their scoring, but I recognize that members are the ones in control of promoting content through their votes.
You are left with what they provide you and can only make small changes to the content on the contact page. To simplify things they even include a contact form that will deliver an email to your inbox, so that visitors don’t need to leave the page to dispatch an email. At the time of this writing I cannot comment any further concerning the ability to sell prints because of this issue. Flickr is great for a layman photographer, who just wants to casually share their photos online, where 500px is meant to be an online showcase of the very best work by professional level photographers.
This problems is partly due to the size of 500px, as it is still a tiny community when compared to Flickr.

I have yet to explore these features because Flickr is overwhelmingly large and offers almost too many features for me to dig through.
SmugMug’s interface is what stops me from creating an account and exploring it further. Even though great photographers are there, it takes hours to find good pictures in Flickr these days. I created some portfolios but can’t for the life of me see how others can view my portfolios.
Most of that traffic is from fellow photographers and those who are interested in viewing beautiful work. These 2 competitors target a much larger audience (artists in general), which is perhaps why 500px has a lead in the photographer market. Photographers earn an 80% commission (minus the cost of print), while Fotomoto will take a 15% cut and 500px a 5% earnings from each sale.
For most photographers this is perfectly acceptable, as they will only submit their top tier work which may consist of only a couple dozen photographs.
500px works best as a social network where you can share your photos with others and receive comments on your work. If you have under 100 great shots that you wish to share you could get away with a basic account if you are patient enough to wait a week between each batch of 20 uploads. I believe that a large amount of photographers on 500px are Russian, so even if they are tagging posts with relevant keywords in their own language, they won’t turn up for the equivalent English search terms.
Because they are targeting a professional level audience they are able to stand out from competitors like Flickr whom appeal to a wide range of styles and skills.
But in smugmug and 500px, I cant see that much number of particular style of photos using their search engine.

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